PhD, biology (ecology), University of Coimbra, 2001.
MSc, biotecnology (biochemistry), IST-Technical University of Lisbon, 1996.
Degree, biology, University of Aveiro, 1992.
Plant systematics and ecology (flora of Portugal, Iberian Peninsula, Angola,Mozambique and East Timor)
Chemotaxonomy
Nature conservation
Environmental impact evaluation and monitoring (flora and vegetation)