PhD in Comparative Biology, Faculty of Philosophy, Sciences and Letters at Ribeirão Preto, University of São Paulo (FFCLRP-USP), Brazil, 2017

Master’s degree in Comparative Biology, Faculty of Philosophy, Sciences and Letters at Ribeirão Preto, University of São Paulo (FFCLRP-USP), Brazil, 2011

Bachelor’s degree in Biology with habilitation in Marine Biology, State University of São Paulo, São Paulo Coast Campus (UNESP-CLP), Brazil, 2008