1. Three-year College course in Biology – 1979
2. Graduate degree in Biology (5 years) – 1981
3. Habilitation for Assistant Lecturer – 1985
4. PhD in Biology – Cell Biology – 1988
5. Habilitation for Full Professorship – 1996
1. Plant Biotechnology
2. Monoterpenes biotransformation by microorganisms
3. Phytochemistry
4. Plant Cell Biology
5. Plant Secretion Biology