2016 – Habilitation (“Agregação”) in Biology, University of Aveiro
1999 – Ph.D. in Biology (Ecology and Biosystematics), University of Lisbon
1993 – M.Sc. in Probability and Statistics, University of Lisbon
1989 – Bachelor (“Licenciatura”) in Biology, University of Lisbon
Photobiology and ecophysiology of marine primary producers.
Photophysiology and productivity of phytoplankton, microphytobenthos, and seagrasses.
Marine photosynthetic symbioses, corals and kleptoplasty.
Photoprotection mechanisms and photoinhibition.
Bioimaging of photosynthesis.
Modelling of carbon fixation and primary production in estuarine environments.