1972 – Graduation in Biology (5-year course, scientific branch), Faculdade de Ciências, Universidade de Lisboa.
1985 – PhD in Biology (Cell Biology), Universidade de Lisboa
Aromatic and Medicinal Plants
Cell Biology
Plant Secretory Structures
Light Microscopy,
Scanning and Transmition Electron Microscopy
Histochemistry and Cytochemistry
Secondary Metabolism
Biological Activities of Natural Products
Nano particles
