1973 – Degree in Biology
1988 – Ph.D. in Biology – Genetics
Eukaryotic viruses: gene expression and replication; studies on antiviral agents.
Viruses from the Portuguese fauna and from water sources: isolation, characterization and molecular identification.
Marine fungi and microalgae: molecular identification and evaluation of activities with potential environmental impact.
Arbuscular Mycorrhizal Fungi (AMF): molecular identification and metagenomics of environmental samples.