1984 – Master of Science in Biology (Biochemistry); Moscow State University named after M.V. Lomonosov, Moscow (U.S.S.R.). Thesis title: [Surface binding and transport of calcium ions in mitochondria]; nº pages: 54; Language: Russian; Supervisor:Iuri Naumovitch Lejkin
1992 – Doutoramento(Ph.D) in Biochemistry; University of Coimbra, Coimbra (Portugal). Thesis title: [Characterization of the transport of the γ-amino-butyric acid in synaptic plasma membrane vesicles]; nº pages: 185; Language: Portuguese; Supervisor: Arsélio Pato de Carvalho
Intercellular communication with focus on nervous and neuroendocrine system. Presynaptic mechanisms of neurotransmission. Role of primary and secondary transport systems in the function of the synapse. Development of experimental models for screening in environmental neurotoxicology.