Licenciatura / First degree in Biology,Universidade de Coimbra, 1984
PhD in Animal and Plant Sciences, University of Sheffield, UK, 1989
Agregação (habilitation for full professorship) in Zoology, Universidade de Coimbra, Portugal, 1997
Ecotoxicologia (solos, dulceaquícola, marinha) / Ecotoxicology (soil, freshwater and marine),
Biodiversidade Funcional / Functional Biodiversity,
Aquacultura / Aquaculture,
Gestão Vida Selvagem / Wildlife Management
Stress